This CARTA symposium addresses the question of how human language came to have the kind of structure it has today, focusing on three sources of evidence. One source, which is discussed in these three talks, concerns neuroscientific investigations of functional specialization for language in the human brain and its dependence on the linguistic input the language learner gets during cognitive development. Evelina Fedorenko (Massachusetts General Hospital) begins with an examination of Specialization for Language in the Human Brain, followed by Rachel Mayberry (UC San Diego) on How the Environment Shapes Language in the Brain, and Edward Chang (UC San Francisco) on Neuroscience of Speech Perception and Speech Production. Recorded on 2/20/2015. (#29395)